When a human body is found inside the strung up Otter mascot uniform of a rival school, Brennan and Booth investigate whether the body inside the costume was dead beforehand, or killed during a Middlesex University bonfire event. The body is identified as Jimmy “Beaver” Bouvier, a fraternity brother at Middlesex University. When Brennan and Booth question his fraternity brothers, they learn that Beaver had a money-making operation on the side and was making a killing of his own. Meanwhile, Booth offers to help his brother Jared find a new job after being dishonorably discharged from the Navy.

A series of indeterminate deaths test the deductive powers of the CSI team, who fear that a public health emergency may be in the offing. Meanwhile, a friendly rivalry over who will win an upcoming bikini contest takes a deadly turn...

As preparations continue for Meredith and Derek's wedding, Izzie receives a surprise hospital visit from her handful-of-a-mother, Robbie. Meanwhile, a newly sober Thatcher visits his daughters, Meredith and Lexie, in order to make amends, and the Chief and Meredith bitterly fight over a controversial medical case.

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Leslie attempts to break into the political "old boy's club," causing an ethical dilemma. Meanwhile, Andy plans a surprise for Ann.

The body Castiel walks around in isn't his. Instead, it belongs to a regular guy named Jimmy who wants his body - and his life - back. He gets the body. But the life may be a different story.

Tonight Stephen welcomes the founder and executive director of the Drug Policy Alliance, Ethan Nadlemann.

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Michael has to mediate a dispute within his new sales team. Meanwhile, trouble brews in the office when several employees take the term "casual" Fridays too loosely.