In Cuba, Nucky presses a U.S. senator on Prohibition's repeal and eyes a business deal with a rum magnate. Margaret witnesses a casualty of the Great Depression. Luciano makes a bold career move. Chalky catches a break to get out of a bad situation. In a flashback to 1884, a young Nucky makes an impression on the Commodore.

Helen and Charlie deal with a crisis. Abby embarks on a journey of self-discovery.

Virginia continues to impersonate a patient's case history in order to glean advice from a psychiatrist. Betty and Masters' attempt to remedy Lester's impotence by hiring a prostitute rattles him even further. Libby gets the third degree when she offers her eyewitness testimony regarding the beaten man. And Barbara Sanders wrestles with an unsettling diagnosis that exposes a chain of secrets and causes Masters and Johnson to clash over their treatment strategy.

Ray tells Bridget how she needs to lie about where she was during the shooting, for her own safety - but Abby brings Jim over to tell her just the opposite. Lee and Ezra are convinced to give Cookie the rights to Marvin and Rekon's music, while Mickey starts to plan a pot store robbery with Shorty. Confronted with Patty's son, Bunchy struggles with his fears. And Kate visits a Walpole inmate to find out what went wrong with her article.

An unexpected ally helps Kevin get out of a jam near the town of Cairo. Meanwhile, an elaborate Memorial Day initiative by the Guilty Remnant plunges the town of Mapleton into chaos. Holy Wayne grants a final wish; Tom copes with his new reality; Nora makes a life-changing decision.

Alison and James share their theory about the source of the fertility crisis with Vanessa. Separately, Vanessa discovers Darius has some co-conspirators as his plans to take over the Lottery continue to escalate. Kyle is asked to compromise his relationship with Allison by the MayTwos and Gabrielle takes the press behind the scenes of The Lottery as contestants face psychological evaluations.

Eph races to safeguard his family, only to learn that Kelly is missing. Dutch reveals her role in Palmer’s plan. Gus is forced to make a difficult decision about Felix.

Part 1 of the three-part premiere of a one-year social experiment involving 15 people who try to create their own civilization from scratch by deciding on a system of government, what food they'll eat and how they'll interact with the outside world.

Freya and Frederick celebrate a milestone birthday. Tarkoff puts pressure on Frederick to get things done while Ingrid and Dash grow closer. Tommy gets a surprise from Wendy as Joanna faces one of her worst fears.